When a former Princeton music professor stumbles into choir practice at a small-town church, he finds singers that are out of tune in more ways than one. Despite their differences, he and his new cohorts may just help each other reinvent.
Arthur defies local customs, and the choir pays for his rudeness. For Cash's sake, Ginny vows to say nothing bad about Wayne.
Arthur helps Ginny conquer stage fright. Reverend Jax is caught in the rift between Wayne and Dwayne.
When Arthur is faced with a choir rebellion, he decides -- against Ginny's advice -- to take on Adams, the group's alpha, at their annual deer hunting trip; Wayne takes Cash on his first hunting trip to teach him how to be a man.
Arthur deals with Jean's father, who attempts to take their family houseboat back. Reverend Jax hosts a singles night, where Ginny and Dwayne finally address the elephant in their relationship.
When Ginny tries to step into a new management role at work, Arthur helps her overcome her fears of inadequacy. When Dwayne opts out of a long-standing Halloween tradition, Wayne fears that they are growing apart.
After Pastor Magnus invites the Second First Choir to perform at the Church of Perpetual Praise, Arthur resolves to sound better than ever to spite him. Ginny spirals when Arthur adds a new singer from her past to the choir.
Reverend Jax asks Arthur to watch the church for him. As parishioners come to him with problems, Arthur believes he can best help them with reason rather than with the ways of the church. Jax also asks Wayne to pick up Adams from the hospital.
Jax convinces Arthur to join the choir's progressive Thanksgiving dinner and snoop in choir members' homes to find a missing item. Meanwhile, Adams invites her handsome contractor to dinner as a set-up for Ginny, but he has something else in mind.
When Reverend Jax is visited by his charismatic parents, Arthur helps him overcome his lifelong feelings of living in their shadow, and encourages him to embrace his way of ministering in time for their Christmas performance. Meanwhile, Ginny and Wayne decide to have a secret fling.
At Conley Fork's annual pageant, judge Arthur tries not to show favoritism toward Cash, in spite of pressure from Ginny. Reverend Jax wonders if Adams should address the unflattering rumors about how her marriage ended.
Arthur gets a surprise visit from a former Princeton colleague, causing him to re-examine his life in Conley Fork. The choir holds an all-night Hymn-A-Thon to raise money for regionals. Ginny attempts to prove that they are not dependent upon Arthur.
At the regional choir competition in Lexington, Arthur relishes the opportunity to finally stick it to his old rival when the unexpected happens. Presented with a surprise job offer, Ginny doubts her worth before her interview. Adams moves on.
